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Bernhard Zipfel

Biographic Data

ID117579
NAMEBernhard Zipfel
GIVEN NAMESBernhard
FAMILY NAMEZipfel
SIGNATUREZIPFEL B
AFFILIATIONSUniversity of the Witwatersrand
ORCID0000-0002-4251-884X
VERIFIEDYes
TOTAL WORKS27
TOTAL CITATIONS106
AUTHOR COUNT26
EDITOR COUNT1
FIRST PUBLICATION YEAR2006
LATEST PUBLICATION YEAR2025
H-INDEX6
  • Catalogue of immature hominin fossils from the University of the Witwatersrand, South Africa

    Open Access•Debra R Bolter, Bernhard Zipfel•ARTICLE•Annals of Human Biology•2025•Cited by: 1•References: 5

    Conclusions: This comprehensive catalogue of immature fossils provides a critical resource for investigating morphological variation, life history traits, and evolutionary adaptations across hominin genera. It underscores the significance of South African fossil collections in exploring developmental patterns and evolutionary pathways leading to the extended life cycle characteristic of Homo sapiens

  • Ecomorphological analysis of bovid remains from the Plio-Pleistocene hominin-bearing deposit of Unit P at Kromdraai, South Africa

    Open Access•Recognise Sambo, R Hanon et al.•ARTICLE•Journal of Archaeological Science…•2024•References: 2

  • An overlooked Australopithecus brain endocast from Makapansgat, South Africa

    Open Access•Z Cofran, Shawn D Hurst et al.•ARTICLE•Journal of Human Evolution•2023•References: 36

  • Les sinus frontaux au cours de l'évolution humaine

    Open Access•Antoine Balzeau, Lou Albessard-Ball et al.•ARTICLE•Cahiers du Centre de recherches…•2023

    Les sinus frontaux sont des cavités qui se situent à l’intérieur de l’os frontal, à la jonction entre le visage et la voûte crânienne, et à proximité du cerveau. Nous proposons l’étude la plus complète jamais réalisée de ces sinus, grâce à une méthodologie facile à mettre en œuvre et reproductible. La taille des sinus frontaux est corrélée à la taille du crâne, et en lien avec celle des reliefs osseux situés au-dessus des orbites, chez les chimpa…

  • The Early Hominin Foot

    Open Access•Jeremy M Desilva, Jeremy Desilva et al.•CHAPTER•Developments in Primatology:…•2022

  • The Kromdraai early hominin-bearing site. A review of recent findings

    Open Access•J Braga, J Francis Thackeray et al.•ARTICLE•L Anthropologie•2022

  • Overlooked or Unimportant? An Overview of the Coprolite Collections at the University of the Witwatersrand, Johannesburg, South Africa

    Open Access•Bernhard Zipfel, C P Montgomery et al.•ARTICLE•Curator The Museum Journal•2022•Cited by: 3•References: 11

    Fossilized feces, termed coprolites, provide unique information on digestive systems, diets, and ecosystems of extinct animals, and are potentially useful for palynology, biostratigraphy and preservation of animal and plant remains. Despite this broad utility, scientific enquiry into coprolites has been relatively sparse. We carried out a systematic investigation into 23 significant South African fossil collections of the Evolutionary Studies Ins…

  • Cochlear morphology of Indonesian Homo erectus from Sangiran

    Open Access•Alessandro Urciuoli, Jülide Kubat et al.•ARTICLE•Journal of Human Evolution•2022•Cited by: 5•References: 77

  • Trabecular bone properties in the ilium of the Middle Paleolithic/Middle Stone Age Border Cave 3 Homo sapiens infant and the onset of independent gait

    Open Access•Kimberleigh A Tommy, Bernhard Zipfel et al.•ARTICLE•Journal of Human Evolution•2021•Cited by: 1•References: 112

  • Hominin Postcranial Remains from Sterkfontein, South Africa, 1936-1995

    Bernhard Zipfel, Brian G Richmond et al.•BOOK•Hominin Postcranial Remains from…•2020

  • Associated Australopithecus afarensis second and third metatarsals (A.L. 333-133) from Hadar, Ethiopia

    Open Access•Jeremy Desilva, Ellison Mcnutt et al.•ARTICLE•Journal of Human Evolution•2020•Cited by: 1•References: 35

  • One small step: A review of Plio‐Pleistocene hominin foot evolution

    Open Access•Jeremy Desilva, Ellison Mcnutt et al.•ARTICLE•American Journal of Physical…•2019

    Bipedalism is a hallmark of being human and the human foot is modified to reflect this unique form of locomotion. Leonardo da Vinci is credited with calling the human foot “a masterpiece of engineering and a work of art.” However, a scientific approach to human origins has revealed that our feet are products of a long, evolutionary history in which a mobile, grasping organ has been converted into a propulsive structure adapted for the rigors of b…

  • Genetic data and radiocarbon dating question Plovers Lake as a Middle Stone Age hominin-bearing site

    Open Access•M Lombard, Helena Malmström et al.•ARTICLE•Journal of Human Evolution•2019•Cited by: 4•References: 20

  • The evolution of the human foot

    Open Access•Ellison Mcnutt, Bernhard Zipfel et al.•ARTICLE•Evolutionary Anthropology Issues…•2018

    There are 26 bones in each foot (52 in total), meaning that roughly a quarter of the human skeleton consists of foot bones. Yet, early hominin foot fossils are frustratingly rare, making it quite difficult to reconstruct the evolutionary history of the human foot. Despite the continued paucity of hominid or hominin foot fossils from the late Miocene and early Pliocene, the last decade has witnessed the discovery of an extraordinary number of earl…

  • Using springbok (Antidorcas) dietary proxies to reconstruct inferred palaeovegetational changes over 2 million years in Southern Africa

    Open Access•Lauren Sewell, Gildas Merceron et al.•ARTICLE•Journal of Archaeological Science…•2018•Cited by: 2•References: 43

  • A quantification of calcaneal lateral plantar process position with implications for bipedal locomotion in Australopithecus

    Open Access•Eve K Boyle, Ellison Mcnutt et al.•ARTICLE•Journal of Human Evolution•2018•Cited by: 10•References: 38

  • Trabecular architecture in the StW 352 fossil hominin calcaneus

    Open Access•Angel Zeininger, B A Patel et al.•ARTICLE•Journal of Human Evolution•2016•Cited by: 18•References: 54

  • The foot of Homo naledi

    Open Access•William E H Harcourt‐smith, W E H Harcourt-Smith et al.•ARTICLE•Nature Communications•2015

    Modern humans are characterized by a highly specialized foot that reflects our obligate bipedalism. Our understanding of hominin foot evolution is, although, hindered by a paucity of well-associated remains. Here we describe the foot of Homo naledi from Dinaledi Chamber, South Africa, using 107 pedal elements, including one nearly-complete adult foot. The H. naledi foot is predominantly modern human-like in morphology and inferred function, with …

  • Recent origin of low trabecular bone density in modern humans

    Open Access•Habiba Chirchir, T L Kivell et al.•ARTICLE•Proceedings of the National…•2015

    Significance The human skeleton is unique in having low trabecular density representing a lightly built human body form. However, it remains unknown when during human evolution this unique characteristic first appeared. To our knowledge, this study is the first to examine trabecular bone density throughout the skeleton of fossil hominins spanning several million years. The results show that trabecular density remained high throughout human evolut…

  • The Lower Limb and Mechanics of Walking in Australopithecus sediba

    Open Access•Jeremy M Desilva, Jeremy Desilva et al.•ARTICLE•Science•2013

    The discovery of a relatively complete Australopithecus sediba adult female skeleton permits a detailed locomotor analysis in which joint systems can be integrated to form a comprehensive picture of gait kinematics in this late australopith. Here we describe the lower limb anatomy of Au. sediba and hypothesize that this species walked with a fully extended leg and with an inverted foot during the swing phase of bipedal walking. Initial contact of…

  • Brief communication: Radiographic study of metatarsal one basal epiphyseal fusion: A note of caution on age determination

    Open Access•E Wei, Elizabeth Weiss et al.•ARTICLE•American Journal of Physical…•2012•Cited by: 2•References: 19

    This study examines radiographs of first metatarsals of 131 individuals from age 17–88 years to determine whether internal basal epiphyseal lines may be visible past the age of metatarsal fusion, which usually occurs between 14 and 16 years of age (Scheuer and Black: The juvenile skeleton. San Diego: Elsevier Academic Press, 2004 ). In 29% (38 out of 131) of the radiographed first metatarsals (MT1s) the basal epiphyseal scar is visible, including…

  • A complete second metatarsal (StW 89) from Sterkfontein Member 4, South Africa

    Open Access•Jeremy M Desilva, Jeremy Desilva et al.•ARTICLE•Journal of Human Evolution•2012•Cited by: 16•References: 40

  • The Foot and Ankle of Australopithecus sediba

    Open Access•Bernhard Zipfel, Jeremy M Desilva et al.•ARTICLE•Science•2011

    Australopithecus sediba had a human-like ankle and arch but an ape-like heel and tibia, implying that while bipedal, this species was also adept at climbing trees.

  • Earliest complete hominin fifth metatarsal: Implications for the evolution of the lateral column of the foot

    Open Access•Bernhard Zipfel, Jeremy Desilva et al.•ARTICLE•HOMO•2010

  • The Olduvai Hominid 8 foot: Adult or subadult

    Open Access•Jeremy M Desilva, Jeremy Desilva et al.•ARTICLE•Journal of Human Evolution•2010•Cited by: 11•References: 22

Next
  • Earliest complete hominin fifth metatarsal—Implications for the evolution of the lateral column of the foot

    Open Access•Bernhard Zipfel, Jeremy M Desilva et al.•ARTICLE•American Journal of Physical…•2009•Cited by: 24•References: 88

    StW 114/115, from Sterkfontein, South Africa, is the earliest complete hominin fifth metatarsal. Comparisons of StW 114/115 to modern humans, extant apes, and partial hominin metatarsals AL 333‐13, AL 333‐78, SKX 33380, OH 8, and KNM‐ER 803f reveal a similar morphology in all six fossils consistent with habitual bipedality. Although StW 114/115 possesses some primitive characters, the proximal articular morphology and internal torsion of the head…

  • Trabecular architecture in the StW 352 fossil hominin calcaneus

    Open Access•Angel Zeininger, B A Patel et al.•ARTICLE•Journal of Human Evolution•2016•Cited by: 18•References: 54

  • A complete second metatarsal (StW 89) from Sterkfontein Member 4, South Africa

    Open Access•Jeremy M Desilva, Jeremy Desilva et al.•ARTICLE•Journal of Human Evolution•2012•Cited by: 16•References: 40

  • The Olduvai Hominid 8 foot: Adult or subadult

    Open Access•Jeremy M Desilva, Jeremy Desilva et al.•ARTICLE•Journal of Human Evolution•2010•Cited by: 11•References: 22

  • A quantification of calcaneal lateral plantar process position with implications for bipedal locomotion in Australopithecus

    Open Access•Eve K Boyle, Ellison Mcnutt et al.•ARTICLE•Journal of Human Evolution•2018•Cited by: 10•References: 38

  • Hominin first metatarsals (SKX 5017 and SK 1813) from Swartkrans: A morphometric analysis

    Open Access•Bernhard Zipfel, Robert S Kidd et al.•ARTICLE•HOMO•2006•Cited by: 8•References: 8

  • Cochlear morphology of Indonesian Homo erectus from Sangiran

    Open Access•Alessandro Urciuoli, Jülide Kubat et al.•ARTICLE•Journal of Human Evolution•2022•Cited by: 5•References: 77

  • Genetic data and radiocarbon dating question Plovers Lake as a Middle Stone Age hominin-bearing site

    Open Access•M Lombard, Helena Malmström et al.•ARTICLE•Journal of Human Evolution•2019•Cited by: 4•References: 20

  • Overlooked or Unimportant? An Overview of the Coprolite Collections at the University of the Witwatersrand, Johannesburg, South Africa

    Open Access•Bernhard Zipfel, C P Montgomery et al.•ARTICLE•Curator The Museum Journal•2022•Cited by: 3•References: 11

    Fossilized feces, termed coprolites, provide unique information on digestive systems, diets, and ecosystems of extinct animals, and are potentially useful for palynology, biostratigraphy and preservation of animal and plant remains. Despite this broad utility, scientific enquiry into coprolites has been relatively sparse. We carried out a systematic investigation into 23 significant South African fossil collections of the Evolutionary Studies Ins…

  • Using springbok (Antidorcas) dietary proxies to reconstruct inferred palaeovegetational changes over 2 million years in Southern Africa

    Open Access•Lauren Sewell, Gildas Merceron et al.•ARTICLE•Journal of Archaeological Science…•2018•Cited by: 2•References: 43

  • Brief communication: Radiographic study of metatarsal one basal epiphyseal fusion: A note of caution on age determination

    Open Access•E Wei, Elizabeth Weiss et al.•ARTICLE•American Journal of Physical…•2012•Cited by: 2•References: 19

    This study examines radiographs of first metatarsals of 131 individuals from age 17–88 years to determine whether internal basal epiphyseal lines may be visible past the age of metatarsal fusion, which usually occurs between 14 and 16 years of age (Scheuer and Black: The juvenile skeleton. San Diego: Elsevier Academic Press, 2004 ). In 29% (38 out of 131) of the radiographed first metatarsals (MT1s) the basal epiphyseal scar is visible, including…

  • Catalogue of immature hominin fossils from the University of the Witwatersrand, South Africa

    Open Access•Debra R Bolter, Bernhard Zipfel•ARTICLE•Annals of Human Biology•2025•Cited by: 1•References: 5

    Conclusions: This comprehensive catalogue of immature fossils provides a critical resource for investigating morphological variation, life history traits, and evolutionary adaptations across hominin genera. It underscores the significance of South African fossil collections in exploring developmental patterns and evolutionary pathways leading to the extended life cycle characteristic of Homo sapiens

  • Trabecular bone properties in the ilium of the Middle Paleolithic/Middle Stone Age Border Cave 3 Homo sapiens infant and the onset of independent gait

    Open Access•Kimberleigh A Tommy, Bernhard Zipfel et al.•ARTICLE•Journal of Human Evolution•2021•Cited by: 1•References: 112

  • Associated Australopithecus afarensis second and third metatarsals (A.L. 333-133) from Hadar, Ethiopia

    Open Access•Jeremy Desilva, Ellison Mcnutt et al.•ARTICLE•Journal of Human Evolution•2020•Cited by: 1•References: 35

  • Hominin first metatarsals (SKX 5017 and SK 1813) from Swartkrans: A morphometric analysis

    Open Access•Bernhard Zipfel, Robert S Kidd et al.•ARTICLE•HOMO•2006•Cited by: 8•References: 8

  • Earliest complete hominin fifth metatarsal—Implications for the evolution of the lateral column of the foot

    Open Access•Bernhard Zipfel, Jeremy M Desilva et al.•ARTICLE•American Journal of Physical…•2009•Cited by: 24•References: 88

    StW 114/115, from Sterkfontein, South Africa, is the earliest complete hominin fifth metatarsal. Comparisons of StW 114/115 to modern humans, extant apes, and partial hominin metatarsals AL 333‐13, AL 333‐78, SKX 33380, OH 8, and KNM‐ER 803f reveal a similar morphology in all six fossils consistent with habitual bipedality. Although StW 114/115 possesses some primitive characters, the proximal articular morphology and internal torsion of the head…

  • Earliest complete hominin fifth metatarsal: Implications for the evolution of the lateral column of the foot

    Open Access•Bernhard Zipfel, Jeremy Desilva et al.•ARTICLE•HOMO•2010

  • The Olduvai Hominid 8 foot: Adult or subadult

    Open Access•Jeremy M Desilva, Jeremy Desilva et al.•ARTICLE•Journal of Human Evolution•2010•Cited by: 11•References: 22

  • The Foot and Ankle of Australopithecus sediba

    Open Access•Bernhard Zipfel, Jeremy M Desilva et al.•ARTICLE•Science•2011

    Australopithecus sediba had a human-like ankle and arch but an ape-like heel and tibia, implying that while bipedal, this species was also adept at climbing trees.

  • Brief communication: Radiographic study of metatarsal one basal epiphyseal fusion: A note of caution on age determination

    Open Access•E Wei, Elizabeth Weiss et al.•ARTICLE•American Journal of Physical…•2012•Cited by: 2•References: 19

    This study examines radiographs of first metatarsals of 131 individuals from age 17–88 years to determine whether internal basal epiphyseal lines may be visible past the age of metatarsal fusion, which usually occurs between 14 and 16 years of age (Scheuer and Black: The juvenile skeleton. San Diego: Elsevier Academic Press, 2004 ). In 29% (38 out of 131) of the radiographed first metatarsals (MT1s) the basal epiphyseal scar is visible, including…

  • A complete second metatarsal (StW 89) from Sterkfontein Member 4, South Africa

    Open Access•Jeremy M Desilva, Jeremy Desilva et al.•ARTICLE•Journal of Human Evolution•2012•Cited by: 16•References: 40

  • The Lower Limb and Mechanics of Walking in Australopithecus sediba

    Open Access•Jeremy M Desilva, Jeremy Desilva et al.•ARTICLE•Science•2013

    The discovery of a relatively complete Australopithecus sediba adult female skeleton permits a detailed locomotor analysis in which joint systems can be integrated to form a comprehensive picture of gait kinematics in this late australopith. Here we describe the lower limb anatomy of Au. sediba and hypothesize that this species walked with a fully extended leg and with an inverted foot during the swing phase of bipedal walking. Initial contact of…

  • The foot of Homo naledi

    Open Access•William E H Harcourt‐smith, W E H Harcourt-Smith et al.•ARTICLE•Nature Communications•2015

    Modern humans are characterized by a highly specialized foot that reflects our obligate bipedalism. Our understanding of hominin foot evolution is, although, hindered by a paucity of well-associated remains. Here we describe the foot of Homo naledi from Dinaledi Chamber, South Africa, using 107 pedal elements, including one nearly-complete adult foot. The H. naledi foot is predominantly modern human-like in morphology and inferred function, with …

  • Recent origin of low trabecular bone density in modern humans

    Open Access•Habiba Chirchir, T L Kivell et al.•ARTICLE•Proceedings of the National…•2015

    Significance The human skeleton is unique in having low trabecular density representing a lightly built human body form. However, it remains unknown when during human evolution this unique characteristic first appeared. To our knowledge, this study is the first to examine trabecular bone density throughout the skeleton of fossil hominins spanning several million years. The results show that trabecular density remained high throughout human evolut…

  • Trabecular architecture in the StW 352 fossil hominin calcaneus

    Open Access•Angel Zeininger, B A Patel et al.•ARTICLE•Journal of Human Evolution•2016•Cited by: 18•References: 54

  • The evolution of the human foot

    Open Access•Ellison Mcnutt, Bernhard Zipfel et al.•ARTICLE•Evolutionary Anthropology Issues…•2018

    There are 26 bones in each foot (52 in total), meaning that roughly a quarter of the human skeleton consists of foot bones. Yet, early hominin foot fossils are frustratingly rare, making it quite difficult to reconstruct the evolutionary history of the human foot. Despite the continued paucity of hominid or hominin foot fossils from the late Miocene and early Pliocene, the last decade has witnessed the discovery of an extraordinary number of earl…

  • Using springbok (Antidorcas) dietary proxies to reconstruct inferred palaeovegetational changes over 2 million years in Southern Africa

    Open Access•Lauren Sewell, Gildas Merceron et al.•ARTICLE•Journal of Archaeological Science…•2018•Cited by: 2•References: 43

  • A quantification of calcaneal lateral plantar process position with implications for bipedal locomotion in Australopithecus

    Open Access•Eve K Boyle, Ellison Mcnutt et al.•ARTICLE•Journal of Human Evolution•2018•Cited by: 10•References: 38

  • One small step: A review of Plio‐Pleistocene hominin foot evolution

    Open Access•Jeremy Desilva, Ellison Mcnutt et al.•ARTICLE•American Journal of Physical…•2019

    Bipedalism is a hallmark of being human and the human foot is modified to reflect this unique form of locomotion. Leonardo da Vinci is credited with calling the human foot “a masterpiece of engineering and a work of art.” However, a scientific approach to human origins has revealed that our feet are products of a long, evolutionary history in which a mobile, grasping organ has been converted into a propulsive structure adapted for the rigors of b…

  • Genetic data and radiocarbon dating question Plovers Lake as a Middle Stone Age hominin-bearing site

    Open Access•M Lombard, Helena Malmström et al.•ARTICLE•Journal of Human Evolution•2019•Cited by: 4•References: 20

  • Hominin Postcranial Remains from Sterkfontein, South Africa, 1936-1995

    Bernhard Zipfel, Brian G Richmond et al.•BOOK•Hominin Postcranial Remains from…•2020

  • Associated Australopithecus afarensis second and third metatarsals (A.L. 333-133) from Hadar, Ethiopia

    Open Access•Jeremy Desilva, Ellison Mcnutt et al.•ARTICLE•Journal of Human Evolution•2020•Cited by: 1•References: 35

  • Trabecular bone properties in the ilium of the Middle Paleolithic/Middle Stone Age Border Cave 3 Homo sapiens infant and the onset of independent gait

    Open Access•Kimberleigh A Tommy, Bernhard Zipfel et al.•ARTICLE•Journal of Human Evolution•2021•Cited by: 1•References: 112

  • The Early Hominin Foot

    Open Access•Jeremy M Desilva, Jeremy Desilva et al.•CHAPTER•Developments in Primatology:…•2022

  • The Kromdraai early hominin-bearing site. A review of recent findings

    Open Access•J Braga, J Francis Thackeray et al.•ARTICLE•L Anthropologie•2022

  • Overlooked or Unimportant? An Overview of the Coprolite Collections at the University of the Witwatersrand, Johannesburg, South Africa

    Open Access•Bernhard Zipfel, C P Montgomery et al.•ARTICLE•Curator The Museum Journal•2022•Cited by: 3•References: 11

    Fossilized feces, termed coprolites, provide unique information on digestive systems, diets, and ecosystems of extinct animals, and are potentially useful for palynology, biostratigraphy and preservation of animal and plant remains. Despite this broad utility, scientific enquiry into coprolites has been relatively sparse. We carried out a systematic investigation into 23 significant South African fossil collections of the Evolutionary Studies Ins…

  • Cochlear morphology of Indonesian Homo erectus from Sangiran

    Open Access•Alessandro Urciuoli, Jülide Kubat et al.•ARTICLE•Journal of Human Evolution•2022•Cited by: 5•References: 77

  • An overlooked Australopithecus brain endocast from Makapansgat, South Africa

    Open Access•Z Cofran, Shawn D Hurst et al.•ARTICLE•Journal of Human Evolution•2023•References: 36

  • Les sinus frontaux au cours de l'évolution humaine

    Open Access•Antoine Balzeau, Lou Albessard-Ball et al.•ARTICLE•Cahiers du Centre de recherches…•2023

    Les sinus frontaux sont des cavités qui se situent à l’intérieur de l’os frontal, à la jonction entre le visage et la voûte crânienne, et à proximité du cerveau. Nous proposons l’étude la plus complète jamais réalisée de ces sinus, grâce à une méthodologie facile à mettre en œuvre et reproductible. La taille des sinus frontaux est corrélée à la taille du crâne, et en lien avec celle des reliefs osseux situés au-dessus des orbites, chez les chimpa…

Biology (22 works) · Pleistocene-Era Hominins and Archaeology (22 works) · Paleontology (17 works) · Evolutionary biology (15 works) · Paleontology (15 works) · Anatomy (14 works) · Evolution and Paleontology Studies (14 works) · Bipedalism (13 works) · Geography (12 works) · Primate Behavior and Ecology (12 works)

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