C Zanolli
Biographic Data
| ID | 27943 |
|---|---|
| NAME | C Zanolli |
| GIVEN NAMES | C |
| FAMILY NAME | Zanolli |
| SIGNATURE | ZANOLLI C |
| AFFILIATIONS | Centre National de la Recherche Scientifique |
| ORCID | 0000-0002-5617-1613 |
| VERIFIED | Yes |
| TOTAL WORKS | 58 |
| TOTAL CITATIONS | 451 |
| AUTHOR COUNT | 58 |
| EDITOR COUNT | 0 |
| FIRST PUBLICATION YEAR | 2010 |
| LATEST PUBLICATION YEAR | 2026 |
| H-INDEX | 12 |
The Early Homo Conundrum: Challenges and Potential Solutions
International audience
Taxonomic revision of the SK 15 mandible based on bone and tooth structural organization
short molar roots) typically found in specimens of Paranthropus aethiopicus, Paranthropus boisei, and Paranthropus robustus. In particular, SK 15 differs markedly in size and morphology from mandibular remains of P. robustus from Swartkrans Member 2. We thus tentatively attribute SK 15 to Paranthropus capensis, a more gracile species of Paranthropus than the other three currently recognized species of this genus and discuss the implications for t…
Étude phénotypique des restes crâniens du site mésolithique de Lake Besaka II (6,8-4,6 ka, Éthiopie)
sont susceptibles d'tre soumis des autorisations d'usage spcifiques
Postcranial evidence does not support habitual bipedalism in Sahelanthropus tchadensis: A reply to Daver et al. (2022)
Une nouvelle approche pour comprendre la robustesse de l’os et de la dentine chez les Néandertaliens: L’étude De La Covariation Entre L’os Cortical Et La Dentine
Une nouvelle approche pour comprendre la robustesse de l'os et de la dentine chez les Nandertaliens : l'tude de la covariation entre l'os cortical et la dentine A new approach to explore bone and dentine robustness in Neandertals: investigation of the covariation between cortical bone and dentine
Enamel thickness in the deciduous postcanine dentition of fossil and extant Pongo
New remains of the Miocene great ape Anoiapithecus brevirostris from Abocador de Can Mata
The dentition of a new adult Neanderthal individual from Grotte Mandrin, France
Grotte Mandrin is located in the middle Rhône River Valley, in Mediterranean France, and has yielded 11 Pleistocene archeological and paleoanthropological layers (ranging from the oldest layer J to the youngest layer B) dating from Marine Isotope Stage (MIS) 5 to MIS 3. We report here the nearly complete dentition of an adult Neanderthal individual, nicknamed 'Thorin,' associated to the last phase of the Post-Neronian II, in layer B2 (∼44.50-42.2…
Three-dimensional geometric morphometric analysis of the bony labyrinth of Xujiayao 6
International audience
Molar enamel-dentine junction shape of Pliobates cataloniae and other Iberian pliopithecoids
Analyse de l'organisation structurale des dents d'un nouvel individu néandertalien de la Grotte Mandrin
La Grotte Mandrin située dans la moyenne vallée du Rhône enregistre 12 niveaux archéologiques pléistocènes (J à B) datés du MIS 5 à la fin du Paléolithique moyen. Nous rapportons ici l’étude de 31 dents permanentes (incluant deux distomolaires) d’un nouvel individu néandertalien adulte surnommé ‘Thorin’ associé au niveau B2 (~42-50 ka) attribué au Post-Néronien II. Une étude paléogénétique a révélé qu’il s’agit d’un individu mâle qui présente une…
Moving away from 'the Muddle in the Middle' toward solving the Chibanian puzzle
Inter- and intrapopulation variability of dental tissue proportions of European and African modern human populations’ permanent canines
International audience
Fifty years of paleoanthropology in Journal of Human Evolution: Historical perspectives and future directions
A reassessment of the distinctiveness of dryopithecine genera from the Iberian Miocene based on enamel-dentine junction geometric morphometric analyses
Estimates of absolute crown strength and bite force in the lower postcanine dentition of Gigantopithecus blacki
Introduction to special issue: Pleistocene hominid diversity and evolution in Asia-A tribute to Pan Lei
Modern human incursion into Neanderthal territories 54,000 years ago at Mandrin, France
Determining the extent of overlap between modern humans and other hominins in Eurasia, such as Neanderthals and Denisovans, is fundamental to understanding the nature of their interactions and what led to the disappearance of archaic hominins. Apart from a possible sporadic pulse recorded in Greece during the Middle Pleistocene, the first settlements of modern humans in Europe have been constrained to ~45,000 to 43,000 years ago. Here, we report …
Dynamics of pre-Hispanic human settlements in the northern Mesoamerican fringe: Contribution of the Study of Variation Based on Tooth Internal Structure
The western Mesoamerica was the place of several human displacements and cultural exchanges. During the Middle Postclassic Period (1200-1450 AD), in the Zacapu basin (located in the NW of the actual state of Michoacán), large urban centers developed, resulting in an increase of the human population. Some hypotheses link this demographic growth to the arrival and the settlement of a new population and likely contributed to the development of these…
Celebrating 50 years of the Journal of Human Evolution
Early Pleistocene hominin teeth from Gongwangling of Lantian, Central China
Further analyses of the structural organization of Homo luzonensis teeth: Evolutionary implications
The species Homo luzonensis has recently been described based on a set of dental and postcranial elements found at Callao Cave (Northern Luzon, Philippines) and dated to at least 50-67 ka. Seven postcanine maxillary teeth are attributed to this taxon, five of them belonging to the same individual (CCH6) and representing the holotype of H. luzonensis, whereas the isolated upper premolar CCH8 and the upper third molar CCH9 are paratypes of the spec…
A deep‐learning‐based workflow to assess taxonomic affinity of hominid teeth with a test on discriminating Pongo and Homo upper molars
OBJECTIVES: Convolutional neural network (CNN) is a state-of-art deep learning (DL) method with superior performance in image classification. Here, a CNN-based workflow is proposed to discriminate hominid teeth. Our hope is that this method could help confirm otherwise questionable records of Homo from Pleistocene deposits where there is a standing risk of mis-attributing molars of Pongo to Homo. METHODS AND MATERIALS: A two-step workflow was des…
A robust alternative to assessing three‐dimensional relative enamel thickness for the use in taxonomic assessment
OBJECTIVE: Three-dimensional relative enamel thickness (3DRET) is important for assessing hypotheses about taxonomy, phylogeny, and dietary reconstruction for primates. However, its weaknesses have not been thoroughly investigated. Here, we analyze its weaknesses and propose an index aiming at better taxonomic discrimination. MATERIALS AND METHODS: The dimensionless 3D index, ratio of enamel-thickness to dentine-thickness (3DRED), which is define…
A comparative analysis of the vestibular apparatus in Epipliopithecus vindobonensis: Phylogenetic implications
Molar crown inner structural organization in J avanese H omo erectus
This contribution investigates the inner organizational pattern (tooth tissue proportions and enamel–dentine junction morphology) of seven Homo erectus permanent molar crowns from the late Lower‐early Middle Pleistocene Kabuh Formation of the Sangiran Dome (Central Java, Indonesia). The previous study of their external characteristics confirmed the degree of time‐related structural reduction occurred in Javanese H. erectus , and also revealed a c…
The late Early Pleistocene human dental remains from Uadi Aalad and Mulhuli-Amo (Buia), Eritrean Danakil: Macromorphology and microstructure
Inner tooth morphology of Homo erectus from Zhoukoudian. New evidence from an old collection housed at Uppsala University, Sweden
Brief communication: Two human fossil deciduous molars from the sangiran dome (Java, Indonesia): Outer and inner morphology
Currently, the human deciduous dental record from the Pleistocene deposits of the Sangiran Dome, Java, consists of only eight specimens. Here we report two deciduous crowns collected near the village of Pucung. While their precise geo‐chronological context remains unknown, a provenance from the Early–Middle Pleistocene Kabuh Formation, or from the Early Pleistocene “Grenzbank Zone,” is very likely. These isolated specimens consist of an upper fir…
A Neanderthal from the Central Western Zagros, Iran. Structural reassessment of the Wezmeh 1 maxillary premolar
Technical note: Morphometric maps of long bone shafts and dental roots for imaging topographic thickness variation
Qualitative and quantitative characterization through functional imaging of mineralized tissues is of potential value in the study of the odontoskeletal remains. This technique, widely developed in the medical field, allows the bi‐dimensional, planar representation of some local morphometric properties, i.e., topographic thickness variation, of a three‐dimensional object, such as a long bone shaft. Nonetheless, the use of morphometric maps is sti…
The early Pleistocene deciduous hominid molar FS ‐72 from the Sangiran dome of java, Indonesia: A taxonomic reappraisal based on its comparative endostructural characterization
Given the morphological and metric similarities between fossil orangutan and H. erectus molars, we tested the hypothesis that its internal morphology more closely resembles the patterns evinced by PCG.2 and modern humans than Pongo. Accordingly, we consider that FS-72 more likely represents a dm2 of Pongo rather than Homo
Intra‐individual metameric variation expressed at the enamel‐dentine junction of lower post‐canine dentition of South African fossil hominins and modern humans
The distalward increase of metameric variation along the modern human molar row is consistent with the odontogenetic models of molar row structure (inhibitory cascade model). Though much remains to be tested, the variable trends and magnitudes in metamerism in fossil hominins reported here, together with differences in the scale of shape change between modern humans and fossil hominins may provide valuable information regarding functional morphol…
Further analyses of the structural organization of Homo luzonensis teeth: Evolutionary implications
The species Homo luzonensis has recently been described based on a set of dental and postcranial elements found at Callao Cave (Northern Luzon, Philippines) and dated to at least 50-67 ka. Seven postcanine maxillary teeth are attributed to this taxon, five of them belonging to the same individual (CCH6) and representing the holotype of H. luzonensis, whereas the isolated upper premolar CCH8 and the upper third molar CCH9 are paratypes of the spec…
New dental remains of Anoiapithecus and the first appearance datum of hominoids in the Iberian Peninsula
Hominin diversity in East Asia during the Middle Pleistocene: A premolar endostructural perspective
Variation in dental morphology and inference of continental ancestry in admixed Latin Americans
Objectives To investigate the variation in dental nonmetric traits and to evaluate the utility of this variation for inferring genetic ancestry proportions in a sample of admixed Latin Americans. Materials and Methods We characterized a sample from Colombia ( N = 477) for 34 dental traits and obtained estimates of individual Native American, European, and African ancestry using genome‐wide SNP data. We tested for correlation between dental traits…
Comparative observations on the premolar root and pulp canal configurations of Middle Pleistocene Homo in China
This work provides new original data, incorporates the latest human fossil discoveries and suggests that analyzing the variation of premolar root structural organization, notably integrating together root/canal form and number, could possibly contribute to taxonomic and phylogenetic assessments. The mid-Middle Pleistocene populations, or "classic" Homo erectus, in our study show closer affinity to Early and Middle Pleistocene hominins in Eurasia,…
Efficacy of diffeomorphic surface matching and 3D geometric morphometrics for taxonomic discrimination of Early Pleistocene hominin mandibular molars
Structural analysis of premolar roots in Middle Pleistocene hominins from China
Contribution of dental tissues to sex determination in modern human populations
OBJECTIVES: Accurate sex estimation is an essential step for the reconstruction of the biological profile of human remains. Earlier studies have shown that elements of the human permanent dentition are sexually dimorphic. The aims of this study are to determine the degree of sexual dimorphism in the dental tissue volumes and surface areas of mandibular canines and to explore its potential for reliable sex determination. METHOD: The teeth included…
A Look from the Inside: MicroCT Analysis of Burned Bones
MicroCT imaging is increasingly used in paleoanthropological and zooarchaeological research to analyse the internal microstructure of bone, replacing comparatively invasive and destructive methods. Consequently the analytical potential of this relatively new 3D imaging technology can be enhanced by developing discipline specific protocols for archaeological analysis. Here we examine how the microstructure of mammal bone changes after burning and …
Dental macrowear and cortical bone distribution of the Neanderthal mandible from Regourdou (Dordogne, Southwestern France)
A comparative analysis of the vestibular apparatus in Epipliopithecus vindobonensis: Phylogenetic implications
A new dryopithecine mandibular fragment from the middle Miocene of Abocador de Can Mata and the taxonomic status of 'Sivapithecus' occidentalis from Can Vila (Vallès-Penedès Basin, NE Iberian Peninsul…
Neutron microtomography‐based virtual extraction and analysis of a cercopithecoid partial cranium ( STS 1039) embedded in a breccia fragment from sterkfontein member 4 (South Africa)
The Plio‐Pleistocene karstic sedimentary deposits of Sterkfontein Cave, South Africa, yielded numerous fossil primate specimens embedded in blocks of indurated breccia, including the partial cercopithecoid cranium labelled STS 1039. Because the surrounding matrix masks most of its morphology, the specimen remains taxonomically undetermined. While the use of X‐ray microtomography did not allow extracting any structural information about the specim…
A reassessment of the distinctiveness of dryopithecine genera from the Iberian Miocene based on enamel-dentine junction geometric morphometric analyses
Modern humans sex estimation through dental tissue patterns of maxillary canines
OBJECTIVES: Dental tissue proportions of human permanent canines is one of only a few sexually dimorphic features that is present in childhood and maintained in adults, offering the opportunity for this to be used in sex determination. This study assesses dental tissue volumes and surface areas of maxillary permanent canines in a sample of known sex to provide new data and to explore the potential of these variables as reliable sexual estimators.…
Stratigraphic context and paleoenvironmental significance of minor taxa (Pisces, Reptilia, Aves, Rodentia) from the late Early Pleistocene paleoanthropological site of Buia (Eritrea)
Moving away from 'the Muddle in the Middle' toward solving the Chibanian puzzle
Technical note: Morphometric maps of long bone shafts and dental roots for imaging topographic thickness variation
Qualitative and quantitative characterization through functional imaging of mineralized tissues is of potential value in the study of the odontoskeletal remains. This technique, widely developed in the medical field, allows the bi‐dimensional, planar representation of some local morphometric properties, i.e., topographic thickness variation, of a three‐dimensional object, such as a long bone shaft. Nonetheless, the use of morphometric maps is sti…
Gestation Length, Mode of Delivery, and Neonatal Line-Thickness Variation
The transition from an intra- to extra-uterine environment leaves its mark in deciduous teeth (and first permanent molars) as an accentuated enamel incremental ring called the neonatal line (NL). This prominent microfeature separates the enamel formed during intrauterine life from that formed after leaving the womb. However, while the physical structure of this scar is well known, the bases of its formation are still a matter of investigation. In…
Brief communication: Two human fossil deciduous molars from the sangiran dome (Java, Indonesia): Outer and inner morphology
Currently, the human deciduous dental record from the Pleistocene deposits of the Sangiran Dome, Java, consists of only eight specimens. Here we report two deciduous crowns collected near the village of Pucung. While their precise geo‐chronological context remains unknown, a provenance from the Early–Middle Pleistocene Kabuh Formation, or from the Early Pleistocene “Grenzbank Zone,” is very likely. These isolated specimens consist of an upper fir…
From outer to inner structural morphology in dental anthropology: Integration of the third dimension in the visualization and quantitative analysis of fossil remains
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New dental remains of Anoiapithecus and the first appearance datum of hominoids in the Iberian Peninsula
Stratigraphic context and paleoenvironmental significance of minor taxa (Pisces, Reptilia, Aves, Rodentia) from the late Early Pleistocene paleoanthropological site of Buia (Eritrea)
The late Early Pleistocene human dental remains from Uadi Aalad and Mulhuli-Amo (Buia), Eritrean Danakil: Macromorphology and microstructure
M icro CT Imaging of Red fox Talus: A Non‐Invasive Approach to Evaluate Age at Death
X‐ray micro CT imaging offers the possibility to study age‐related changes of bone microstructure. In the present paper we analyse the talus of 15 modern red foxes of different ages, from 2 months old to adulthood, to investigate the possibility of identifying their different ages at death. Surface and volumetric variables describing bone properties are measured or evaluated from the micro CT images following three approaches: (i) the bone volume…
Molar crown inner structural organization in J avanese H omo erectus
This contribution investigates the inner organizational pattern (tooth tissue proportions and enamel–dentine junction morphology) of seven Homo erectus permanent molar crowns from the late Lower‐early Middle Pleistocene Kabuh Formation of the Sangiran Dome (Central Java, Indonesia). The previous study of their external characteristics confirmed the degree of time‐related structural reduction occurred in Javanese H. erectus , and also revealed a c…
The early Pleistocene deciduous hominid molar FS ‐72 from the Sangiran dome of java, Indonesia: A taxonomic reappraisal based on its comparative endostructural characterization
Given the morphological and metric similarities between fossil orangutan and H. erectus molars, we tested the hypothesis that its internal morphology more closely resembles the patterns evinced by PCG.2 and modern humans than Pongo. Accordingly, we consider that FS-72 more likely represents a dm2 of Pongo rather than Homo
A Look from the Inside: MicroCT Analysis of Burned Bones
MicroCT imaging is increasingly used in paleoanthropological and zooarchaeological research to analyse the internal microstructure of bone, replacing comparatively invasive and destructive methods. Consequently the analytical potential of this relatively new 3D imaging technology can be enhanced by developing discipline specific protocols for archaeological analysis. Here we examine how the microstructure of mammal bone changes after burning and …
Neutron microtomography‐based virtual extraction and analysis of a cercopithecoid partial cranium ( STS 1039) embedded in a breccia fragment from sterkfontein member 4 (South Africa)
The Plio‐Pleistocene karstic sedimentary deposits of Sterkfontein Cave, South Africa, yielded numerous fossil primate specimens embedded in blocks of indurated breccia, including the partial cercopithecoid cranium labelled STS 1039. Because the surrounding matrix masks most of its morphology, the specimen remains taxonomically undetermined. While the use of X‐ray microtomography did not allow extracting any structural information about the specim…
Intra‐individual metameric variation expressed at the enamel‐dentine junction of lower post‐canine dentition of South African fossil hominins and modern humans
The distalward increase of metameric variation along the modern human molar row is consistent with the odontogenetic models of molar row structure (inhibitory cascade model). Though much remains to be tested, the variable trends and magnitudes in metamerism in fossil hominins reported here, together with differences in the scale of shape change between modern humans and fossil hominins may provide valuable information regarding functional morphol…
Structural organization and tooth development in a Homo aff. erectus juvenile mandible from the Early Pleistocene site of Garba IV at Melka Kunture, Ethiopian highlands
OBJECTIVES: The immature partial mandible GAR IVE from the c. 1.7 Ma old Garba IV site at Melka Kunture (Upper Awash Basin, Ethiopia), the earliest human representative from a mountain-like environment, represents one of the oldest early Homo specimens bearing a mixed dentition. Following its first description (Condemi, ), we extended the analytical and comparative record of this specimen by providing unreported details about its inner morphology…
Contribution of dental tissues to sex determination in modern human populations
OBJECTIVES: Accurate sex estimation is an essential step for the reconstruction of the biological profile of human remains. Earlier studies have shown that elements of the human permanent dentition are sexually dimorphic. The aims of this study are to determine the degree of sexual dimorphism in the dental tissue volumes and surface areas of mandibular canines and to explore its potential for reliable sex determination. METHOD: The teeth included…
Neutron‐based computed microtomography: Pliobates cataloniae and Barberapithecus huerzeleri as a test‐case study
OBJECTIVES: High-resolution imaging of fossils with X-ray computed microtomography (μCT) has become a very powerful tool in paleontological research. However, fossilized bone, embedding matrix, and dental tissues do not always provide a distinct structural signal with X-rays. We demonstrate the benefits of high-resolution neutron radiation in three different specimens showing problematic contrasts with X-ray μCT. MATERIALS AND METHODS: We compare…
Modern humans sex estimation through dental tissue patterns of maxillary canines
OBJECTIVES: Dental tissue proportions of human permanent canines is one of only a few sexually dimorphic features that is present in childhood and maintained in adults, offering the opportunity for this to be used in sex determination. This study assesses dental tissue volumes and surface areas of maxillary permanent canines in a sample of known sex to provide new data and to explore the potential of these variables as reliable sexual estimators.…
Inner tooth morphology of Homo erectus from Zhoukoudian. New evidence from an old collection housed at Uppsala University, Sweden
A new species of Homo from the Late Pleistocene of the Philippines
A Neanderthal from the Central Western Zagros, Iran. Structural reassessment of the Wezmeh 1 maxillary premolar
Comparative observations on the premolar root and pulp canal configurations of Middle Pleistocene Homo in China
This work provides new original data, incorporates the latest human fossil discoveries and suggests that analyzing the variation of premolar root structural organization, notably integrating together root/canal form and number, could possibly contribute to taxonomic and phylogenetic assessments. The mid-Middle Pleistocene populations, or "classic" Homo erectus, in our study show closer affinity to Early and Middle Pleistocene hominins in Eurasia,…
A distinguishing feature of Pongo upper molars and its implications for the taxonomic identification of isolated hominid teeth from the Pleistocene of Asia
This study suggests that the protoconule and the shape of the mesial fovea in upper molars are useful features for the taxonomic identification of isolated hominid teeth. By identifying these new features, our results can contribute to the better understanding of hominoid evolutionary history and biogeography during the Asian Pleistocene. However, we emphasize that the reported features should be used in combination with other diagnostic variable…
Variation in dental morphology and inference of continental ancestry in admixed Latin Americans
Objectives To investigate the variation in dental nonmetric traits and to evaluate the utility of this variation for inferring genetic ancestry proportions in a sample of admixed Latin Americans. Materials and Methods We characterized a sample from Colombia ( N = 477) for 34 dental traits and obtained estimates of individual Native American, European, and African ancestry using genome‐wide SNP data. We tested for correlation between dental traits…
A geometric morphometric approach to the study of variation of shovel‐shaped incisors
OBJECTIVES: The scoring and analysis of dental nonmetric traits are predominantly accomplished by using the Arizona State University Dental Anthropology System (ASUDAS), a standard protocol based on strict definitions and three-dimensional dental plaques. However, visual scoring, even when controlled by strict definitions of features, visual reference, and the experience of the observer, includes an unavoidable part of subjectivity. In this metho…
Efficacy of diffeomorphic surface matching and 3D geometric morphometrics for taxonomic discrimination of Early Pleistocene hominin mandibular molars
Biology (47 works) · Pleistocene-Era Hominins and Archaeology (46 works) · Forensic Anthropology and Bioarchaeology Studies (37 works) · Paleontology (34 works) · Paleontology (33 works) · Evolution and Paleontology Studies (27 works) · Geology (26 works) · Geology (26 works) · Evolutionary biology (22 works) · Geography (22 works)