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Grant Ramsey

Biographic Data

ID3866155
NAMEGrant Ramsey
GIVEN NAMESGrant
FAMILY NAMERamsey
SIGNATURERAMSEY G
AFFILIATIONSKU Leuven
ORCID0000-0002-8712-5521
VERIFIEDYes
TOTAL WORKS21
TOTAL CITATIONS97
WORKS AS AUTHOR20
WORKS AS EDITOR1
FIRST PUBLICATION YEAR2006
LATEST PUBLICATION YEAR2025
H-INDEX7
  • Tool Use Beyond Humans

    Open Access•Gianmaria Dani, Grant Ramsey•ARTICLE•Philosophy of Science•2025•References: 40

    The definition of tool use has long been debated, especially when applied beyond humans. Recent work argues that the phenomena included within tool use are so broad and varied that there is little hope of using the category for scientific generalizations, explanations, and predictions about the evolution, ecology, and psychology of tool users. One response to this argument has been the development of tooling as a replacement for tool use. In this…

  • What Are the ‘Levels’ in Levels of Selection

    Markus I Eronen, Grant Ramsey•ARTICLE•The British Journal for the…•2025•References: 55

    The levels of selection debate is generally taken to be a debate about how natural selection can occur at the various levels of biological organization.In this paper, we argue that questions about levels of selection should be analyzed separately from questions about levels of organization.In the deflationary proposal we defend, all that is necessary for multilevel selection is that there are cases in which particles are nested in collectives, an…

  • Driftability and niche construction

    Open Access•Alejandro Fábregas-Tejeda, Grant Ramsey•ARTICLE•Synthese•2024•References: 84

    Niche construction is the process of organisms changing themselves or their environment—or their relationship with their environment—in ways that affect the evolutionary trajectory of their population. These evolutionary trajectory changes are traditionally understood to be triggered by changes in selection pressures. Niche construction thus necessarily involves organisms altering selection pressures. In this paper, we argue that changes in selec…

  • Developmental Channeling and Evolutionary Dappling

    Open Access•Grant Ramsey, Cristina Villegas•ARTICLE•Philosophy of Science•2024•References: 53

    The developmental properties of organisms play important roles in the generation of variation necessary for evolutionary change. But how can individual development steer the course of evolution? To answer this question, we introduce developmental channeling as a disposition of individual organisms that shapes their possible developmental trajectories and evolutionary dappling as an evolutionary outcome in which the space of possible organismic fo…

  • The concepts and origins of cell mortality

    Open Access•Pierre Durand, Pierre M Durand et al.•ARTICLE•History & Philosophy of the Life…•2023

    Organismal death is foundational to the evolution of life, and many biological concepts such as natural selection and life history strategy are so fashioned only because individuals are mortal. Organisms, irrespective of their organization, are composed of basic functional units-cells-and it is our understanding of cell death that lies at the heart of most general explanatory frameworks for organismal mortality. Cell death can be exogenous, arisi…

  • Adaptationism and Trait Individuation

    Open Access•James Difrisco, Grant Ramsey•ARTICLE•Philosophy of Science•2023•Cited by: 2•References: 19

    Adaptationism is often taken to be the thesis that most traits are adaptations. To assess this thesis, it seems we must be able to establish either an exhaustive set of all traits or a representative sample of this set. Either task requires a more systematic and principled way of individuating traits than is currently available. Moreover, different trait individuation criteria can make adaptationism turn out true or false. For instance, individua…

  • Human Success

    Hugh Desmond, Grant Ramsey•EDITOR•BOOK•Human Success•2023

  • Empathy and the Evolutionary Emergence of Guilt

    Open Access•Grant Ramsey, Michael J Deem•ARTICLE•Philosophy of Science•2022•Cited by: 2•References: 56

    Guilt poses a unique evolutionary problem. Unlike other dysphoric emotions, it is not immediately clear what its adaptive significance is. One can imagine thriving despite or even because of a lack of guilt. In this article, we review solutions offered by Scott James, Richard Joyce, and Robert Frank and show that although their solutions have merit, none adequately solves the puzzle. We offer an alternative solution, one that emphasizes the role …

  • Three Kinds of Niche Construction

    Open Access•Bendik Hellem Aaby, Grant Ramsey•ARTICLE•The British Journal for the…•2022•Cited by: 9•References: 52

    Niche construction theory concerns how organisms can change selection pressures by altering the feature–factor relationship between themselves and their environment. These alterations are standardly understood to be brought about through two kinds of organism–environment interaction: perturbative and relocational niche construction. We argue that a reconceptualization is needed on the grounds that if a niche is understood as the feature–factor re…

  • How to Do Digital Philosophy of Science

    Open Access•Charles H Pence, Grant Ramsey•ARTICLE•Philosophy of Science•2018•Cited by: 12•References: 21

    Philosophy of science is expanding via the introduction of new digital data and tools for their analysis. The data comprise digitized published books and journal articles, as well as heretofore unpublished material such as images, archival text, notebooks, meeting notes, and programs. The growth in available data is matched by the extensive development of automated analysis tools. The variety of data sources and tools can be overwhelming. In this…

  • Is Cultural Fitness Hopelessly Confused

    Grant Ramsey, Andreas De Block•ARTICLE•The British Journal for the…•2017•Cited by: 3•References: 37

    Fitness is a central concept in evolutionary theory. Just as it is central to biological evolution, so, it seems, it should be central to cultural evolutionary theory (CET). But importing the biological fitness concept to CET is no straightforward task—there are many features unique to cultural evolution that make this difficult. This has led some theorists to argue that there are fundamental problems with cultural fitness that render it hopeless…

  • EvoText

    Open Access•Grant Ramsey, Charles H Pence•ARTICLE•Studies in History and Philosophy…•2016•Cited by: 4•References: 1

    We introduce here evoText, a new tool for automated analysis of the literature in the biological sciences. evoText contains a database of hundreds of thousands of journal articles and an array of analysis tools for generating quantitative data on the nature and history of life science, especially ecology and evolutionary biology. This article describes the features of evoText, presents a variety of examples of the kinds of analyses that evoText c…

  • Can altruism be unified

    Open Access•Grant Ramsey•ARTICLE•Studies in History and Philosophy…•2015•Cited by: 2•References: 2

  • Is Organismic Fitness at the Basis of Evolutionary Theory

    Open Access•Charles H Pence, Grant Ramsey•ARTICLE•Philosophy of Science•2015•Cited by: 4•References: 10

    Fitness is a central theoretical concept in evolutionary theory. Despite its importance, much debate has occurred over how to conceptualize and formalize fitness. One point of debate concerns the roles of organismic and trait fitness. In a recent addition to this debate, Elliott Sober argues that trait fitness is the central fitness concept, and that organismic fitness is of little value. In this paper, by contrast, we argue that it is organismic…

  • Driftability

    Open Access•Grant Ramsey•ARTICLE•Synthese•2013•Cited by: 9•References: 24

  • Human Nature in a Post-essentialist World

    Open Access•Grant Ramsey•ARTICLE•Philosophy of Science•2013•Cited by: 7•References: 4

    In this essay I examine a well-known articulation of human nature skepticism, a paper by Hull. I then review a recent reply to Hull by Machery, which argues for an account of human nature that he claims is both useful and scientifically robust. I challenge Machery’s account and introduce an alternative account—the “life-history trait cluster” conception of human nature—that I hold is scientifically sound and makes sense of (at least some of) our …

  • Organisms, Traits, and Population Subdivisions

    Grant Ramsey•ARTICLE•The British Journal for the…•2013•Cited by: 6•References: 26

    A major debate in the philosophy of biology centers on the question of how we should understand the causal structure of natural selection. This debate is polarized into the causal and statistical positions. The main arguments from the statistical side are that a causal construal of the theory of natural selection's central concept, fitness, either (i) leads to inaccurate predictions about population dynamics, or (ii) leads to an incoherent set of…

  • A New Foundation for the Propensity Interpretation of Fitness

    Charles H Pence, Grant Ramsey•ARTICLE•The British Journal for the…•2013•Cited by: 19•References: 56

    The propensity interpretation of fitness (PIF) is commonly taken to be subject to a set of simple counterexamples. We argue that three of the most important of these are not counterexamples to the PIF itself, but only to the traditional mathematical model of this propensity: fitness as expected number of offspring. They fail to demonstrate that a new mathematical model of the PIF could not succeed where this older model fails. We then propose a n…

  • How Human Nature Can Inform Human Enhancement

    Open Access•Grant Ramsey•ARTICLE•Philosophy & Technology•2012

  • Sameness in Biology

    Open Access•Grant Ramsey, Anne Siebels Peterson•ARTICLE•Philosophy of Science•2012•Cited by: 7•References: 17

    Homology is a biological sameness relation that is purported to hold in the face of changes in form, composition, and function. In spite of the centrality and importance of homology, there is no consensus on how we should understand this concept. The two leading views of homology, the genealogical and developmental accounts, have significant shortcomings. We propose a new account, the hierarchical-dependency account of homology, which avoids thes…

  • Block Fitness

    Open Access•Grant Ramsey•ARTICLE•Studies in History and Philosophy…•2006•Cited by: 11

  • A New Foundation for the Propensity Interpretation of Fitness

    Charles H Pence, Grant Ramsey•ARTICLE•The British Journal for the…•2013•Cited by: 19•References: 56

    The propensity interpretation of fitness (PIF) is commonly taken to be subject to a set of simple counterexamples. We argue that three of the most important of these are not counterexamples to the PIF itself, but only to the traditional mathematical model of this propensity: fitness as expected number of offspring. They fail to demonstrate that a new mathematical model of the PIF could not succeed where this older model fails. We then propose a n…

  • How to Do Digital Philosophy of Science

    Open Access•Charles H Pence, Grant Ramsey•ARTICLE•Philosophy of Science•2018•Cited by: 12•References: 21

    Philosophy of science is expanding via the introduction of new digital data and tools for their analysis. The data comprise digitized published books and journal articles, as well as heretofore unpublished material such as images, archival text, notebooks, meeting notes, and programs. The growth in available data is matched by the extensive development of automated analysis tools. The variety of data sources and tools can be overwhelming. In this…

  • Block Fitness

    Open Access•Grant Ramsey•ARTICLE•Studies in History and Philosophy…•2006•Cited by: 11

  • Three Kinds of Niche Construction

    Open Access•Bendik Hellem Aaby, Grant Ramsey•ARTICLE•The British Journal for the…•2022•Cited by: 9•References: 52

    Niche construction theory concerns how organisms can change selection pressures by altering the feature–factor relationship between themselves and their environment. These alterations are standardly understood to be brought about through two kinds of organism–environment interaction: perturbative and relocational niche construction. We argue that a reconceptualization is needed on the grounds that if a niche is understood as the feature–factor re…

  • Driftability

    Open Access•Grant Ramsey•ARTICLE•Synthese•2013•Cited by: 9•References: 24

  • Human Nature in a Post-essentialist World

    Open Access•Grant Ramsey•ARTICLE•Philosophy of Science•2013•Cited by: 7•References: 4

    In this essay I examine a well-known articulation of human nature skepticism, a paper by Hull. I then review a recent reply to Hull by Machery, which argues for an account of human nature that he claims is both useful and scientifically robust. I challenge Machery’s account and introduce an alternative account—the “life-history trait cluster” conception of human nature—that I hold is scientifically sound and makes sense of (at least some of) our …

  • Sameness in Biology

    Open Access•Grant Ramsey, Anne Siebels Peterson•ARTICLE•Philosophy of Science•2012•Cited by: 7•References: 17

    Homology is a biological sameness relation that is purported to hold in the face of changes in form, composition, and function. In spite of the centrality and importance of homology, there is no consensus on how we should understand this concept. The two leading views of homology, the genealogical and developmental accounts, have significant shortcomings. We propose a new account, the hierarchical-dependency account of homology, which avoids thes…

  • Organisms, Traits, and Population Subdivisions

    Grant Ramsey•ARTICLE•The British Journal for the…•2013•Cited by: 6•References: 26

    A major debate in the philosophy of biology centers on the question of how we should understand the causal structure of natural selection. This debate is polarized into the causal and statistical positions. The main arguments from the statistical side are that a causal construal of the theory of natural selection's central concept, fitness, either (i) leads to inaccurate predictions about population dynamics, or (ii) leads to an incoherent set of…

  • EvoText

    Open Access•Grant Ramsey, Charles H Pence•ARTICLE•Studies in History and Philosophy…•2016•Cited by: 4•References: 1

    We introduce here evoText, a new tool for automated analysis of the literature in the biological sciences. evoText contains a database of hundreds of thousands of journal articles and an array of analysis tools for generating quantitative data on the nature and history of life science, especially ecology and evolutionary biology. This article describes the features of evoText, presents a variety of examples of the kinds of analyses that evoText c…

  • Is Organismic Fitness at the Basis of Evolutionary Theory

    Open Access•Charles H Pence, Grant Ramsey•ARTICLE•Philosophy of Science•2015•Cited by: 4•References: 10

    Fitness is a central theoretical concept in evolutionary theory. Despite its importance, much debate has occurred over how to conceptualize and formalize fitness. One point of debate concerns the roles of organismic and trait fitness. In a recent addition to this debate, Elliott Sober argues that trait fitness is the central fitness concept, and that organismic fitness is of little value. In this paper, by contrast, we argue that it is organismic…

  • Is Cultural Fitness Hopelessly Confused

    Grant Ramsey, Andreas De Block•ARTICLE•The British Journal for the…•2017•Cited by: 3•References: 37

    Fitness is a central concept in evolutionary theory. Just as it is central to biological evolution, so, it seems, it should be central to cultural evolutionary theory (CET). But importing the biological fitness concept to CET is no straightforward task—there are many features unique to cultural evolution that make this difficult. This has led some theorists to argue that there are fundamental problems with cultural fitness that render it hopeless…

  • Adaptationism and Trait Individuation

    Open Access•James Difrisco, Grant Ramsey•ARTICLE•Philosophy of Science•2023•Cited by: 2•References: 19

    Adaptationism is often taken to be the thesis that most traits are adaptations. To assess this thesis, it seems we must be able to establish either an exhaustive set of all traits or a representative sample of this set. Either task requires a more systematic and principled way of individuating traits than is currently available. Moreover, different trait individuation criteria can make adaptationism turn out true or false. For instance, individua…

  • Empathy and the Evolutionary Emergence of Guilt

    Open Access•Grant Ramsey, Michael J Deem•ARTICLE•Philosophy of Science•2022•Cited by: 2•References: 56

    Guilt poses a unique evolutionary problem. Unlike other dysphoric emotions, it is not immediately clear what its adaptive significance is. One can imagine thriving despite or even because of a lack of guilt. In this article, we review solutions offered by Scott James, Richard Joyce, and Robert Frank and show that although their solutions have merit, none adequately solves the puzzle. We offer an alternative solution, one that emphasizes the role …

  • Can altruism be unified

    Open Access•Grant Ramsey•ARTICLE•Studies in History and Philosophy…•2015•Cited by: 2•References: 2

  • Block Fitness

    Open Access•Grant Ramsey•ARTICLE•Studies in History and Philosophy…•2006•Cited by: 11

  • How Human Nature Can Inform Human Enhancement

    Open Access•Grant Ramsey•ARTICLE•Philosophy & Technology•2012

  • Sameness in Biology

    Open Access•Grant Ramsey, Anne Siebels Peterson•ARTICLE•Philosophy of Science•2012•Cited by: 7•References: 17

    Homology is a biological sameness relation that is purported to hold in the face of changes in form, composition, and function. In spite of the centrality and importance of homology, there is no consensus on how we should understand this concept. The two leading views of homology, the genealogical and developmental accounts, have significant shortcomings. We propose a new account, the hierarchical-dependency account of homology, which avoids thes…

  • Driftability

    Open Access•Grant Ramsey•ARTICLE•Synthese•2013•Cited by: 9•References: 24

  • Human Nature in a Post-essentialist World

    Open Access•Grant Ramsey•ARTICLE•Philosophy of Science•2013•Cited by: 7•References: 4

    In this essay I examine a well-known articulation of human nature skepticism, a paper by Hull. I then review a recent reply to Hull by Machery, which argues for an account of human nature that he claims is both useful and scientifically robust. I challenge Machery’s account and introduce an alternative account—the “life-history trait cluster” conception of human nature—that I hold is scientifically sound and makes sense of (at least some of) our …

  • Organisms, Traits, and Population Subdivisions

    Grant Ramsey•ARTICLE•The British Journal for the…•2013•Cited by: 6•References: 26

    A major debate in the philosophy of biology centers on the question of how we should understand the causal structure of natural selection. This debate is polarized into the causal and statistical positions. The main arguments from the statistical side are that a causal construal of the theory of natural selection's central concept, fitness, either (i) leads to inaccurate predictions about population dynamics, or (ii) leads to an incoherent set of…

  • A New Foundation for the Propensity Interpretation of Fitness

    Charles H Pence, Grant Ramsey•ARTICLE•The British Journal for the…•2013•Cited by: 19•References: 56

    The propensity interpretation of fitness (PIF) is commonly taken to be subject to a set of simple counterexamples. We argue that three of the most important of these are not counterexamples to the PIF itself, but only to the traditional mathematical model of this propensity: fitness as expected number of offspring. They fail to demonstrate that a new mathematical model of the PIF could not succeed where this older model fails. We then propose a n…

  • Can altruism be unified

    Open Access•Grant Ramsey•ARTICLE•Studies in History and Philosophy…•2015•Cited by: 2•References: 2

  • Is Organismic Fitness at the Basis of Evolutionary Theory

    Open Access•Charles H Pence, Grant Ramsey•ARTICLE•Philosophy of Science•2015•Cited by: 4•References: 10

    Fitness is a central theoretical concept in evolutionary theory. Despite its importance, much debate has occurred over how to conceptualize and formalize fitness. One point of debate concerns the roles of organismic and trait fitness. In a recent addition to this debate, Elliott Sober argues that trait fitness is the central fitness concept, and that organismic fitness is of little value. In this paper, by contrast, we argue that it is organismic…

  • EvoText

    Open Access•Grant Ramsey, Charles H Pence•ARTICLE•Studies in History and Philosophy…•2016•Cited by: 4•References: 1

    We introduce here evoText, a new tool for automated analysis of the literature in the biological sciences. evoText contains a database of hundreds of thousands of journal articles and an array of analysis tools for generating quantitative data on the nature and history of life science, especially ecology and evolutionary biology. This article describes the features of evoText, presents a variety of examples of the kinds of analyses that evoText c…

  • Is Cultural Fitness Hopelessly Confused

    Grant Ramsey, Andreas De Block•ARTICLE•The British Journal for the…•2017•Cited by: 3•References: 37

    Fitness is a central concept in evolutionary theory. Just as it is central to biological evolution, so, it seems, it should be central to cultural evolutionary theory (CET). But importing the biological fitness concept to CET is no straightforward task—there are many features unique to cultural evolution that make this difficult. This has led some theorists to argue that there are fundamental problems with cultural fitness that render it hopeless…

  • How to Do Digital Philosophy of Science

    Open Access•Charles H Pence, Grant Ramsey•ARTICLE•Philosophy of Science•2018•Cited by: 12•References: 21

    Philosophy of science is expanding via the introduction of new digital data and tools for their analysis. The data comprise digitized published books and journal articles, as well as heretofore unpublished material such as images, archival text, notebooks, meeting notes, and programs. The growth in available data is matched by the extensive development of automated analysis tools. The variety of data sources and tools can be overwhelming. In this…

  • Empathy and the Evolutionary Emergence of Guilt

    Open Access•Grant Ramsey, Michael J Deem•ARTICLE•Philosophy of Science•2022•Cited by: 2•References: 56

    Guilt poses a unique evolutionary problem. Unlike other dysphoric emotions, it is not immediately clear what its adaptive significance is. One can imagine thriving despite or even because of a lack of guilt. In this article, we review solutions offered by Scott James, Richard Joyce, and Robert Frank and show that although their solutions have merit, none adequately solves the puzzle. We offer an alternative solution, one that emphasizes the role …

  • Three Kinds of Niche Construction

    Open Access•Bendik Hellem Aaby, Grant Ramsey•ARTICLE•The British Journal for the…•2022•Cited by: 9•References: 52

    Niche construction theory concerns how organisms can change selection pressures by altering the feature–factor relationship between themselves and their environment. These alterations are standardly understood to be brought about through two kinds of organism–environment interaction: perturbative and relocational niche construction. We argue that a reconceptualization is needed on the grounds that if a niche is understood as the feature–factor re…

  • The concepts and origins of cell mortality

    Open Access•Pierre Durand, Pierre M Durand et al.•ARTICLE•History & Philosophy of the Life…•2023

    Organismal death is foundational to the evolution of life, and many biological concepts such as natural selection and life history strategy are so fashioned only because individuals are mortal. Organisms, irrespective of their organization, are composed of basic functional units-cells-and it is our understanding of cell death that lies at the heart of most general explanatory frameworks for organismal mortality. Cell death can be exogenous, arisi…

  • Adaptationism and Trait Individuation

    Open Access•James Difrisco, Grant Ramsey•ARTICLE•Philosophy of Science•2023•Cited by: 2•References: 19

    Adaptationism is often taken to be the thesis that most traits are adaptations. To assess this thesis, it seems we must be able to establish either an exhaustive set of all traits or a representative sample of this set. Either task requires a more systematic and principled way of individuating traits than is currently available. Moreover, different trait individuation criteria can make adaptationism turn out true or false. For instance, individua…

  • Human Success

    Hugh Desmond, Grant Ramsey•EDITOR•BOOK•Human Success•2023

  • Driftability and niche construction

    Open Access•Alejandro Fábregas-Tejeda, Grant Ramsey•ARTICLE•Synthese•2024•References: 84

    Niche construction is the process of organisms changing themselves or their environment—or their relationship with their environment—in ways that affect the evolutionary trajectory of their population. These evolutionary trajectory changes are traditionally understood to be triggered by changes in selection pressures. Niche construction thus necessarily involves organisms altering selection pressures. In this paper, we argue that changes in selec…

  • Developmental Channeling and Evolutionary Dappling

    Open Access•Grant Ramsey, Cristina Villegas•ARTICLE•Philosophy of Science•2024•References: 53

    The developmental properties of organisms play important roles in the generation of variation necessary for evolutionary change. But how can individual development steer the course of evolution? To answer this question, we introduce developmental channeling as a disposition of individual organisms that shapes their possible developmental trajectories and evolutionary dappling as an evolutionary outcome in which the space of possible organismic fo…

  • Tool Use Beyond Humans

    Open Access•Gianmaria Dani, Grant Ramsey•ARTICLE•Philosophy of Science•2025•References: 40

    The definition of tool use has long been debated, especially when applied beyond humans. Recent work argues that the phenomena included within tool use are so broad and varied that there is little hope of using the category for scientific generalizations, explanations, and predictions about the evolution, ecology, and psychology of tool users. One response to this argument has been the development of tooling as a replacement for tool use. In this…

  • What Are the ‘Levels’ in Levels of Selection

    Markus I Eronen, Grant Ramsey•ARTICLE•The British Journal for the…•2025•References: 55

    The levels of selection debate is generally taken to be a debate about how natural selection can occur at the various levels of biological organization.In this paper, we argue that questions about levels of selection should be analyzed separately from questions about levels of organization.In the deflationary proposal we defend, all that is necessary for multilevel selection is that there are cases in which particles are nested in collectives, an…

Philosophy (19 works) · Epistemology (18 works) · Biology (11 works) · Computer Science (11 works) · Evolutionary Game Theory and Cooperation (11 works) · Sociology (11 works) · Evolution and Genetic Dynamics (10 works) · Philosophy and History of Science (9 works) · Psychology (9 works) · Artificial Intelligence (6 works)

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