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First Population Assessment Report, Suitable Habitat and Ecological Corridors of the Endangered Kashmir Gray Langur Morphotype ( Semnopithecus ajax , Pocock 1928) in Khyber Pakhtunkhwa, Pakistan

Bibliographic Data

ID19388650
AuthorsAbid Ali (0000-0002-0452-4827, Chinese Academy of Sciences, corresponding author), David Walter Rössel‐Ramírez (0000-0002-1056-9992, Technological University of Mexico), Arif Jan (0000-0002-2768-3986, Oregon Department of Fish and Wildlife), Idris Sari (Erzincan Binali Yıldırım University), Adrián Martín-Taboada (0000-0002-8269-5090, Biogeography, Diversity, and Conservation Research Team, Department of Animal Biology, Faculty of Sciences University of Malaga Malaga Spain), Adrián Martín‐Taboada (Universidad de Málaga), Jorge Juan‐Rueda (0009-0009-3051-9894, University of Alicante), Milenka Djukanovich‐Iturbe (0000-0003-2833-0768, Universidad Nacional de Córdoba), Iftikhar U-Zaman (0000-0001-8385-2353, Climate Change, Forestry Environment, and Wildlife Department Government of Khyber Pakhtunkhwa Peshawar Pakistan), José Vicente López‐bao (0000-0001-9213-998X, Biodiversity Research Institute), Muhammad Noman Khan (0000-0001-6475-8126, Quaid-i-Azam University)
Year2026
Volume88
Issue1
Pagese70117-e70117
Publication date2026-01-01
Peer ReviewedYes
Open AccessYes
TypeARTICLE
VenueAmerican Journal of Primatology (JOURNAL)
Journal identifiersISSN: 0275-2565 • E-ISSN: 1098-2345
PublisherWiley (PUBLISHER • GB)
DOI10.1002/ajp.70117
PMID41548199
OpenAlexW7124703653
LanguageEN
References cited62

The Himalayan Kashmir langur ( Semnopithecus ajax ) is an endangered colobine of the northwestern Himalayas, occurring in India and Pakistan; yet its ecology remains poorly studied. In Pakistan, it is known from Azad Jammu and Kashmir (AJK) and the northern districts of Khyber Pakhtunkhwa (KPK) but populations restricted to KPK has been neglected until the data. In this regard, we conducted 60 line‐transect surveys (118.2 km) across 69.5 km2 in three KPK districts—Mansehra, Kohistan, and Battagram—to estimate population density and abundance. Further, we modeled habitat suitability and connectivity modeling of langurs across its known range in Pakistan. The current study revealed mean density across the surveyed districts was 4.5 individuals/km2 (95% CI: 2.3–6.7), with substantially higher densities in Mansehra (7.4; 95% CI: 3.3–11.5) compared to Kohistan–Battagram (KB) (1.3; 95% CI: 0.1–2.5). The total population in the study area was estimated at 314 individuals (95% CI: 143–485), with ~86% concentrated in Mansehra. Habitat suitability modeling (AUC = 0.96) identified four major suitable niche zones (SNZ) (~2,953 km2) across Pakistan, primarily montane forests between 1,300–3,100 m with > 40% canopy cover, though only ~82 km2 (~1%) overlapped with existing protected areas. Connectivity analysis indicated forested corridors linking these niche zones, while fragmentation reduced connectivity in several regions. Our findings demonstrate that the langur persists in Pakistan as a small, fragmented, and forest‐dependent population. Conservation of this threatened species will require improved forest protection, management of habitat corridors, and coordinated transboundary conservation efforts across its Himalayan range

Conservation status · Endangered species · Habitat · Habitat destruction · National park · Population · Population density · Population size · Range (aeronautics) · Threatened species · Amphibian and Reptile Biology · Species Distribution and Climate Change · Wildlife Ecology and Conservation

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